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Recent Classifications

Four recent classifications of the Anthocerotes

Hässel de Menéndez (1988)

Anthocerotales Limpricht in Cohn
    Anthocerotaceae Dum.
    Anthoceros (Mich.) L.
    Spaerosporoceros Hässel
    Notothyladaceae (Milde) Müll.
        Subfam. Notothyladoideae Grolle
               Notothylas Sull.
        Subfam. Phaeocerotoideae Hässel
            Phaeoceros Prosk.
Foliocerotales Hässel de Menéndez
    Foliocerotaceae Hässel
    Folioceros Bharadwaj
Leiosporocerotales Hässel de Menéndez
    Leiosporocerotaceae Hässel de Menéndez
    Leiosporoceros Hässel de Men.
Dendrocerotales Hässel de Menéndez
    Dendrocerotaceae (Milde) Hässel de Men.
    Dendroceros Nees
    Megaceros Campbell

Schuster (1992)

Anthocerotales Limpricht in Cohn
     Anthocerotaceae Dum.
        Subfam. Anthocerotoideae Dumort.
              *Aspiromitus Steph. emend. Rink
                  Subgen. Aspiromitus
                  Subgen. Folioceros
                      *Anthoceros (Mich.) L.
                  Subgen. Anthoceros
                  Subgen. Leiosporoceros
         Subfam. Dendrocerotoideae Schuster
               Megaceros Campbell
                   Subgen. Megaceros
                   Subgen. Notoceros Schuster
                         Dendroceros Nees
                   Subgen. Dendroceros
                   Subgen. Apoceros Schuster
          Subfam. Notothyladoideae Grolle
                            Notothylas Sull.
* This author recognizes Aspiromitus in place of Anthoceros and
Anthoceros in place of Phaeoceros

Hyvönen and Piippo (1993)

Anthocerotales Limpricht in Cohn
Anthocerotaceae Dum.
Anthoceros (Mich.) L.
Folioceros Bharadwaj
Leiosporoceros Hässel de Men.
Mesoceros Piippo
Phaeoceros Prosk.
Spaerosporoceros Hässel de Men.
Dendrocerotaceae (Milde) Hässel de Men.
Dendroceros Nees
Megaceros Campbell
Nothothylales
Notothlyadaceae (Milde) Müll.
Notothylas Sull.

Hasegawa (1994)

Anthocerotales Limpricht in Cohn
       Notothyladaceae (Milde) Müll.
            Notothylas Sull.
       Anthocerotaceae Dum.
            Subfam. Anthocerotoideae
                   Anthoceros (Mich.) L.
                   Subgen. Anthoceros
                   Subgen. Folioceros
                   Leiosporoceros Hässel de Men.
                   Hattorioceros Hasegawa
                   Phaeoceros Prosk.
              Subfam. Dendrocerotoideae
                  Dendroceros Nees
                  Subgen. Dendroceros
                  Subgen. Apoceros Schuster
                      Megaceros Campbell
                      Notoceros (Schuster) Hasegawa

References:

Hässel de Menéndez, G. G. 1988. A proposal for a new classification of the genera within the Anthocerotophyta. J. Hattori Bot. Lab 64: 71-86. As pointed out by Hyvönen and Piippo (1993), some nodes on this tree (shown with dashed lines) have no character support and are unresolved.

Shuster, ??????

Hyvönen, J. and S. Piippo. 1993. Cladistic analysis of the hornworts (Anthocerotophyta). J. Hattori Bot. Lab 74: 105-119.

Hasegawa, J. 1994. New classification of Anthocerotae. J. Hattori Bot. Lab 76: 21-34.